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    The Origin of Species

    Page 31
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    3000 feet. Prof. Ramsay has published an account of a downthrow in

      Anglesea of 2300 feet; and he informs me that he fully believes there is

      one in Merionethshire of 12,000 feet; yet in these cases there is nothing

      on the surface to show such prodigious movements; the pile of rocks on the

      one or other side having been smoothly swept away. The consideration of

      these facts impresses my mind almost in the same manner as does the vain

      endeavour to grapple with the idea of eternity.

      I am tempted to give one other case, the well-known one of the denudation

      of the Weald. Though it must be admitted that the denudation of the Weald

      has been a mere trifle, in comparison with that which has removed masses of

      our palaeozoic strata, in parts ten thousand feet in thickness, as shown in

      Prof. Ramsay's masterly memoir on this subject. Yet it is an admirable

      lesson to stand on the North Downs and to look at the distant South Downs;

      for, remembering that at no great distance to the west the northern and

      southern escarpments meet and close, one can safely picture to oneself the

      great dome of rocks which must have covered up the Weald within so limited

      a period as since the latter part of the Chalk formation. The distance

      from the northern to the southern Downs is about 22 miles, and the

      thickness of the several formations is on an average about 1100 feet, as I

      am informed by Prof. Ramsay. But if, as some geologists suppose, a range

      of older rocks underlies the Weald, on the flanks of which the overlying

      sedimentary deposits might have accumulated in thinner masses than

      elsewhere, the above estimate would be erroneous; but this source of doubt

      probably would not greatly affect the estimate as applied to the western

      extremity of the district. If, then, we knew the rate at which the sea

      commonly wears away a line of cliff of any given height, we could measure

      the time requisite to have denuded the Weald. This, of course, cannot be

      done; but we may, in order to form some crude notion on the subject, assume

      that the sea would eat into cliffs 500 feet in height at the rate of one

      inch in a century. This will at first appear much too small an allowance;

      but it is the same as if we were to assume a cliff one yard in height to be

      eaten back along a whole line of coast at the rate of one yard in nearly

      every twenty-two years. I doubt whether any rock, even as soft as chalk,

      would yield at this rate excepting on the most exposed coasts; though no

      doubt the degradation of a lofty cliff would be more rapid from the

      breakage of the fallen fragments. On the other hand, I do not believe that

      any line of coast, ten or twenty miles in length, ever suffers degradation

      at the same time along its whole indented length; and we must remember that

      almost all strata contain harder layers or nodules, which from long

      resisting attrition form a breakwater at the base. Hence, under ordinary

      circumstances, I conclude that for a cliff 500 feet in height, a denudation

      of one inch per century for the whole length would be an ample allowance.

      At this rate, on the above data, the denudation of the Weald must have

      required 306,662,400 years; or say three hundred million years.

      The action of fresh water on the gently inclined Wealden district, when

      upraised, could hardly have been great, but it would somewhat reduce the

      above estimate. On the other hand, during oscillations of level, which we

      know this area has undergone, the surface may have existed for millions of

      years as land, and thus have escaped the action of the sea: when deeply

      submerged for perhaps equally long periods, it would, likewise, have

      escaped the action of the coast-waves. So that in all probability a far

      longer period than 300 million years has elapsed since the latter part of

      the Secondary period.

      I have made these few remarks because it is highly important for us to gain

      some notion, however imperfect, of the lapse of years. During each of

      these years, over the whole world, the land and the water has been peopled

      by hosts of living forms. What an infinite number of generations, which

      the mind cannot grasp, must have succeeded each other in the long roll of

      years! Now turn to our richest geological museums, and what a paltry

      display we behold!

      On the poorness of our Palaeontological collections. -- That our

      palaeontological collections are very imperfect, is admitted by every one.

      The remark of that admirable palaeontologist, the late Edward Forbes,

      should not be forgotten, namely, that numbers of our fossil species are

      known and named from single and often broken specimens, or from a few

      specimens collected on some one spot. Only a small portion of the surface

      of the earth has been geologically explored, and no part with sufficient

      care, as the important discoveries made every year in Europe prove. No

      organism wholly soft can be preserved. Shells and bones will decay and

      disappear when left on the bottom of the sea, where sediment is not

      accumulating. I believe we are continually taking a most erroneous view,

      when we tacitly admit to ourselves that sediment is being deposited over

      nearly the whole bed of the sea, at a rate sufficiently quick to embed and

      preserve fossil remains. Throughout an enormously large proportion of the

      ocean, the bright blue tint of the water bespeaks its purity. The many

      cases on record of a formation conformably covered, after an enormous

      interval of time, by another and later formation, without the underlying

      bed having suffered in the interval any wear and tear, seem explicable only

      on the view of the bottom of the sea not rarely lying for ages in an

      unaltered condition. The remains which do become embedded, if in sand or

      gravel, will when the beds are upraised generally be dissolved by the

      percolation of rain-water. I suspect that but few of the very many animals

      which live on the beach between high and low watermark are preserved. For

      instance, the several species of the Chthamalinae (a sub-family of sessile

      cirripedes) coat the rocks all over the world in infinite numbers: they

      are all strictly littoral, with the exception of a single Mediterranean

      species, which inhabits deep water and has been found fossil in Sicily,

      whereas not one other species has hitherto been found in any tertiary

      formation: yet it is now known that the genus Chthamalus existed during

      the chalk period. The molluscan genus Chiton offers a partially analogous

      case.

      With respect to the terrestrial productions which lived during the

      Secondary and Palaeozoic periods, it is superfluous to state that our

      evidence from fossil remains is fragmentary in an extreme degree. For

      instance, not a land shell is known belonging to either of these vast

      periods, with one exception discovered by Sir C. Lyell in the carboniferous

      strata of North America. In regard to mammiferous remains, a single glance

      at the historical table published in the Supplement to Lyell's Manual, will

      bring home the truth, how accidental and rare is their preservation, far

      better than pages of detail. Nor is their rarity surprising, when we

      remember how
    large a proportion of the bones of tertiary mammals have been

      discovered either in caves or in lacustrine deposits; and that not a cave

      or true lacustrine bed is known belonging to the age of our secondary or

      palaeozoic formations.

      But the imperfection in the geological record mainly results from another

      and more important cause than any of the foregoing; namely, from the

      several formations being separated from each other by wide intervals of

      time. When we see the formations tabulated in written works, or when we

      follow them in nature, it is difficult to avoid believing that they are

      closely consecutive. But we know, for instance, from Sir R. Murchison's

      great work on Russia, what wide gaps there are in that country between the

      superimposed formations; so it is in North America, and in many other parts

      of the world. The most skilful geologist, if his attention had been

      exclusively confined to these large territories, would never have suspected

      that during the periods which were blank and barren in his own country,

      great piles of sediment, charged with new and peculiar forms of life, had

      elsewhere been accumulated. And if in each separate territory, hardly any

      idea can be formed of the length of time which has elapsed between the

      consecutive formations, we may infer that this could nowhere be

      ascertained. The frequent and great changes in the mineralogical

      composition of consecutive formations, generally implying great changes in

      the geography of the surrounding lands, whence the sediment has been

      derived, accords with the belief of vast intervals of time having elapsed

      between each formation.

      But we can, I think, see why the geological formations of each region are

      almost invariably intermittent; that is, have not followed each other in

      close sequence. Scarcely any fact struck me more when examining many

      hundred miles of the South American coasts, which have been upraised

      several hundred feet within the recent period, than the absence of any

      recent deposits sufficiently extensive to last for even a short geological

      period. Along the whole west coast, which is inhabited by a peculiar

      marine fauna, tertiary beds are so scantily developed, that no record of

      several successive and peculiar marine faunas will probably be preserved to

      a distant age. A little reflection will explain why along the rising coast

      of the western side of South America, no extensive formations with recent

      or tertiary remains can anywhere be found, though the supply of sediment

      must for ages have been great, from the enormous degradation of the

      coast-rocks and from muddy streams entering the sea. The explanation, no

      doubt, is, that the littoral and sub-littoral deposits are continually worn

      away, as soon as they are brought up by the slow and gradual rising of the

      land within the grinding action of the coast-waves.

      We may, I think, safely conclude that sediment must be accumulated in

      extremely thick, solid, or extensive masses, in order to withstand the

      incessant action of the waves, when first upraised and during subsequent

      oscillations of level. Such thick and extensive accumulations of sediment

      may be formed in two ways; either, in profound depths of the sea, in which

      case, judging from the researches of E. Forbes, we may conclude that the

      bottom will be inhabited by extremely few animals, and the mass when

      upraised will give a most imperfect record of the forms of life which then

      existed; or, sediment may be accumulated to any thickness and extent over a

      shallow bottom, if it continue slowly to subside. In this latter case, as

      long as the rate of subsidence and supply of sediment nearly balance each

      other, the sea will remain shallow and favourable for life, and thus a

      fossiliferous formation thick enough, when upraised, to resist any amount

      of degradation, may be formed.

      I am convinced that all our ancient formations, which are rich in fossils,

      have thus been formed during subsidence. Since publishing my views on this

      subject in 1845, I have watched the progress of Geology, and have been

      surprised to note how author after author, in treating of this or that

      great formation, has come to the conclusion that it was accumulated during

      subsidence. I may add, that the only ancient tertiary formation on the

      west coast of South America, which has been bulky enough to resist such

      degradation as it has as yet suffered, but which will hardly last to a

      distant geological age, was certainly deposited during a downward

      oscillation of level, and thus gained considerable thickness.

      All geological facts tell us plainly that each area has undergone numerous

      slow oscillations of level, and apparently these oscillations have affected

      wide spaces. Consequently formations rich in fossils and sufficiently

      thick and extensive to resist subsequent degradation, may have been formed

      over wide spaces during periods of subsidence, but only where the supply of

      sediment was sufficient to keep the sea shallow and to embed and preserve

      the remains before they had time to decay. On the other hand, as long as

      the bed of the sea remained stationary, thick deposits could not have been

      accumulated in the shallow parts, which are the most favourable to life.

      Still less could this have happened during the alternate periods of

      elevation; or, to speak more accurately, the beds which were then

      accumulated will have been destroyed by being upraised and brought within

      the limits of the coast-action.

      Thus the geological record will almost necessarily be rendered

      intermittent. I feel much confidence in the truth of these views, for they

      are in strict accordance with the general principles inculcated by Sir C.

      Lyell; and E. Forbes independently arrived at a similar conclusion.

      One remark is here worth a passing notice. During periods of elevation the

      area of the land and of the adjoining shoal parts of the sea will be

      increased, and new stations will often be formed;--all circumstances most

      favourable, as previously explained, for the formation of new varieties and

      species; but during such periods there will generally be a blank in the

      geological record. On the other hand, during subsidence, the inhabited

      area and number of inhabitants will decrease (excepting the productions on

      the shores of a continent when first broken up into an archipelago), and

      consequently during subsidence, though there will be much extinction, fewer

      new varieties or species will be formed; and it is during these very

      periods of subsidence, that our great deposits rich in fossils have been

      accumulated. Nature may almost be said to have guarded against the

      frequent discovery of her transitional or linking forms.

      From the foregoing considerations it cannot be doubted that the geological

      record, viewed as a whole, is extremely imperfect; but if we confine our

      attention to any one formation, it becomes more difficult to understand,

      why we do not therein find closely graduated varieties between the allied

      species which lived at its commencement and at its close. Some cases are

      on record of the same species presenting distinct varieties in the upper


      and lower parts of the same formation, but, as they are rare, they may be

      here passed over. Although each formation has indisputably required a vast

      number of years for its deposition, I can see several reasons why each

      should not include a graduated series of links between the species which

      then lived; but I can by no means pretend to assign due proportional weight

      to the following considerations.

      Although each formation may mark a very long lapse of years, each perhaps

      is short compared with the period requisite to change one species into

      another. I am aware that two palaeontologists, whose opinions are worthy

      of much deference, namely Bronn and Woodward, have concluded that the

      average duration of each formation is twice or thrice as long as the

      average duration of specific forms. But insuperable difficulties, as it

      seems to me, prevent us coming to any just conclusion on this head. When

      we see a species first appearing in the middle of any formation, it would

      be rash in the extreme to infer that it had not elsewhere previously

      existed. So again when we find a species disappearing before the uppermost

      layers have been deposited, it would be equally rash to suppose that it

      then became wholly extinct. We forget how small the area of Europe is

      compared with the rest of the world; nor have the several stages of the

      same formation throughout Europe been correlated with perfect accuracy.

      With marine animals of all kinds, we may safely infer a large amount of

      migration during climatal and other changes; and when we see a species

      first appearing in any formation, the probability is that it only then

      first immigrated into that area. It is well known, for instance, that

      several species appeared somewhat earlier in the palaeozoic beds of North

      America than in those of Europe; time having apparently been required for

      their migration from the American to the European seas. In examining the

      latest deposits of various quarters of the world, it has everywhere been

      noted, that some few still existing species are common in the deposit, but

      have become extinct in the immediately surrounding sea; or, conversely,

      that some are now abundant in the neighbouring sea, but are rare or absent

      in this particular deposit. It is an excellent lesson to reflect on the

      ascertained amount of migration of the inhabitants of Europe during the

      Glacial period, which forms only a part of one whole geological period; and

      likewise to reflect on the great changes of level, on the inordinately

      great change of climate, on the prodigious lapse of time, all included

      within this same glacial period. Yet it may be doubted whether in any

      quarter of the world, sedimentary deposits, including fossil remains, have

      gone on accumulating within the same area during the whole of this period.

      It is not, for instance, probable that sediment was deposited during the

      whole of the glacial period near the mouth of the Mississippi, within that

      limit of depth at which marine animals can flourish; for we know what vast

      geographical changes occurred in other parts of America during this space

      of time. When such beds as were deposited in shallow water near the mouth

      of the Mississippi during some part of the glacial period shall have been

      upraised, organic remains will probably first appear and disappear at

      different levels, owing to the migration of species and to geographical

      changes. And in the distant future, a geologist examining these beds,

      might be tempted to conclude that the average duration of life of the

      embedded fossils had been less than that of the glacial period, instead of

      having been really far greater, that is extending from before the glacial

      epoch to the present day.

      In order to get a perfect gradation between two forms in the upper and

      lower parts of the same formation, the deposit must have gone on

      accumulating for a very long period, in order to have given sufficient time

     


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